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‘Bees are sentient’: inside the stunning brains of nature’s hardest workers
(www.theguardian.com)
My totally uneducated guess: bees have evolved to be even more attuned to the environment than many other insects. They’re aware of changes that we can’t detect (or haven’t thought to look for). They can sense their environment collapsing. They’re “unhappy” because they can tell that things aren’t right.